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... that it functioned far better using  spike timing 
Adapting a Feedforward Heteroassociative Network to Hodgkin-Huxley Dynamics
Adapting a Feedforward Heteroassociative Network to Hodgkin-Huxley Dynamics
Lytton, William W.
, rather than spike occurrence, as the ...
... preserve information encoded in precise  spike timing 
Cholinergic Modulation of Spike Timing and Spike Frequency Adaptation in Neocortical Neurons
Cholinergic Modulation of Spike Timing and Spike Frequency Adaptation in Neocortical Neurons
Tang, A. C.; Bartels, A. M.; Sejnowski, T. J.
, but not in interspike intervals, and ...