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... the cells not signal stimulus polarity.  Spike timing 
The temporal properties of non-linear, transient cells in the locust medulla
The temporal properties of non-linear, transient cells in the locust medulla
Osorio, D.
depends upon stimulus contrast, below ...
... elevated, with a characteristic post-  spike time 
Activity-related changes in electrical thresholds of pyramidal tract axons in the behaving monkey
Activity-related changes in electrical thresholds of pyramidal tract axons in the behaving monkey
Schmied, A.; Fetz, E. E.
course. ...